Swap the tubes, one type at a time, from one channel to the other. When the whine moves to the other channel you'll know the tubes you just changed were the culprits. Start the with single 12AX7, then the pairs of 12AU7 then the pairs of KT88 (move these types as pairs only since they are likely matched and should stay that way). If the whine never does move to the other channel then something else is amiss. I think that amp is self-biasing. If it isn't, than rebias after each swap.