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Depends on how the designer designed the speaker. Mine sounds better to me without the grill cloth, but I have heard speakers that need a pillow over them to tame the top end. I would leave the grill cloth on with those, for sure, and maybe add an extra towel too.
I have some Salk Songtowers (Ribbon Tweeter version) and definitely hear the difference with grills off and the owner (Jim Salk) tells you to remove them for critical listening. they are magnetic and easy to remove and put back on. I typically leave them on more for protection when not in use or just watching TV. Figure at least might help with a flying dog toy or something.
The difference in sound isn't so much from the grill cloth, but from the diffraction off the frame of the grill.
Say NO 2 the cloth...to be naked is to BE.