Replacing an 5AR4 with a 5R4GY in an amplifier designed for a 5AR4 simply lowers the B+ voltage, turns down the output tubes, may make the bias adjustment go out of range low, lowers the power capability, provides a higher then design load on the filament circuit, and increases the distortion.
Not recommended with an Ultravalve or Ultimate 70, although some may like the sonic results with other designs. Changing tube types without an understanding of the circuit or making measurements is like testing tires without a pressure gauge.
Installing an appropriate diode in series with each lead of the transformer B+ supply will unload the 5AR4, using it essentially just for its slow turn on effect and maintaining the appropriate voltage drop for design B+ voltage under load and improve 5AR4 service life. Recommended. Standard on Ultravalve amps.
Frank Van Alstine