I have been listening to the 550exr with the rev5 upgrade for a couple days now. IMO, and in my system, it is an improvement.
The highs are "more" and sweeter. The cymbals have more shimmer and decay -- and are more natural sounding imo. The low volume level "background" highs( micro highs ? ) are now more hearable --- before i thought they were slightly attenuated. So overall there is a tad more detail.
The low frequencies are still great, but maybe have a little more "clarity" .
The mids are too tough a call imo.
Please understand that these comments are based on "aural memory", and not a direct comparison between two amps. YMMV
Hope this might help someone that is considering this upgrade..........