Brad,
After this work, I'm doubting that the problem lies with the tubes or the sockets.
It could be the grid coupling cap. These caps, if overheated during soldering, will eventually fail, giving the symptoms you describe.
Polystyrene caps are highly sensitive to heat, and their failure mode is erratic, and in this case they have about 45 volts across them, so they are electrically stressed.
My suggestion would be to replace it with a high quality filmcap, and see if that fixes it.
Hope this helps,
Hugh